Many times when you see a weird assortment of actions with lights and gauges, it's a lack of voltage issue. Dead batts and/or connections.
And with that, you had a no-crank issue.
The dreaded fuel limp;
Many things can be the problem.
There's a lot left out of the story/time line. Did you pull over and stop to clear the code? Just trying to figure out why a restart (that didn't) occured.
1.5 hrs pulling a wind load, if you started with a full tank, you were probably down to 1/2-5/8 tank level. And as you drive under load, you're also returning hot fuel to the slowly dropping fuel tank levels....continually raising the fuel temps. Hotter fuel can trigger seal issues.
Tank fuel module could be a cause at lower tank levels (cracked/leaking/sucking air).
Moving forward, you've already addressed possible collapsed fuel lines. Next is the filter and filter head. How old is the filter? Is it still the original filter head, or has it been changed yet?
Clogged filter can kick the fuel limp also.
Was that trip starting tank purchased from a different source than you usually use?
Was it higher in bio levels than what's normally used?
Fuel sourced could have prematurely clogged the fuel filter. Especially if it was a higher bio level (solvent) combined with higher fuel temps.
The fuel cooler for the returning fuel doesn't do much. A full tank of fresh cool fuel is the main heatsink. Long trips in the heat of the day, towing a heavy wind load, you'll see higher return fuel temps dumping into a shrinking level of what was once cool fuel.
Hotter fuel affects everything in the fuel system and can trigger problems with components/seals. Many times, just stopping and topping off the tank with fresh cool fuel can settle everything down.....for a while.
Higher than normal fuel return rates from an injector(s) or FPRV can quickly raise temps too.
I'd start with a fresh filter, unless you have a gauge to measure at the filter head, psi @idle to determine any restriction level,
Voltage; load test your batts, independently. Inspect connections/cable conditions, as well as all the main harness connections.
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